Malagasy civet

Species of carnivore

Malagasy civet
at Ranomafana National Park
Conservation status

Vulnerable  (IUCN 3.1)[1]
CITES Appendix II (CITES)[2]
Scientific classification Edit this classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Mammalia
Order: Carnivora
Suborder: Feliformia
Family: Eupleridae
Subfamily: Euplerinae
Genus: Fossa
Gray, 1864[3]
Species:
F. fossana
Binomial name
Fossa fossana
(Müller, 1776)[3]
Malagasy civet range[4]
Synonyms

Viverra fossana Müller, 1776

The Malagasy or striped civet (Fossa fossana), also known as the fanaloka (Malagasy, [fə̥ˈnaluk]) or jabady,[5] is an euplerid endemic to Madagascar.[6] It is the only species in genus Fossa.

The Malagasy civet is a small mammal, about 47 centimetres (19 in) long excluding the tail (which is only about 20 centimetres (7.9 in)). It can weigh 1.5 to 2.0 kilograms (3.3 to 4.4 lb). It is endemic to the tropical forests of Madagascar. Malagasy civets are nocturnal. It eats small vertebrates, insects, aquatic animals, and eggs stolen from birds' nests. The mating season of the Malagasy civet is August to September and the gestation period is three months, ending with the birth of one young. The Malagasy civet is listed as Vulnerable by International Union for Conservation of Nature (IUCN).[1]

The Malagasy name fanaloka is related to the Malay word pelanduk "mousedeer" (via metathesis) likely due to their similar sizes and silhouette.[7]

Classification

The Malagasy civet was to be placed in the subfamily Hemigalinae with the banded palm civets and then in its own subfamily, Fossinae, because of similarities with others in the group pointed out by Gregory, but it is now classified as a member of the subfamily Euplerinae, after Pocock pointed out more similarities with that one.[8]

Description

The Malagasy civet is a small mammal, about 47 centimetres (19 in) long excluding the tail (which is only about 20 centimetres (7.9 in)). The males can weigh up to 1.9 kilograms (4.2 lb), and the females can weigh up to 1.75 kilograms (3.9 lb). It is the second largest carnivore in Madagascar after the fossa.[9][10] it may be confused with the small Indian civet (Viverricula indica).[11] It has a short coat greyish beige or brown in colour, with dark black horizontal stripes running from head to tail, where the stripes are vertical, wrapping around the bushier tail. The stripes morph into spots near the belly. Its legs are short and very thin.[11]

Behavior

The Malagasy civet is nocturnal, though sources disagree over whether it is solitary or, unusual among euplerids, lives in pairs. It is not a good climber and frequents ravines. It eats small vertebrates (mammals, small birds, reptiles, and amphibians), insects, aquatic animals, and eggs stolen from birds' nests.[12][13] It is shy and secretive.[14] Their vocalizations are similar to crying and groaning, as well as a sound similar to coq-coq. Pairs of males and females defend a large area (around 50 hectares (120 acres)) as their territory.[15] In the winter, it may store fat in its tail, which can make up 25% of their weight.[9][16] The mating season of the Malagasy civet is August to September and the gestation period is around three months, ending with the birth of one young. The young are rather well-developed, weigh around 65 to 70 grams (2.3 to 2.5 oz), and are weaned in two to three months, leaving their parents at around one year old. The average lifespan of a Malagasy civet is about 21 years in captivity.[1][17][18]

Distribution and habitat

The Malagasy civet is found in lowland and rainforest areas of Eastern and Northern areas of Madagascar, and can also be found in humid and isolated forests in Amber Mountain National Park, and farther north in the less-humid forests of Ankarana Reserve. It can be found from sea level to 1,600 metres (5,200 ft) above sea level, but is only common up to 1,000 metres (3,300 ft) above sea level.[1][11]

Conservation status

The Malagasy civet is listed as Vulnerable by International Union for Conservation of Nature (IUCN), with a decreasing population. Though threatened by deforestation, hunting, charcoal production, logging, and competition from introduced species such as dogs, cats, and small Indian civets, it is locally common. Introduced animals such as dogs are likely to prey on Malagasy civets.[1] Its range is now reduced to isolated patches.[18]

References

  1. ^ a b c d e Hawkins, F. (2015). "Fossa fossana". IUCN Red List of Threatened Species. 2015: e.T8668A45197868. doi:10.2305/IUCN.UK.2015-4.RLTS.T8668A45197868.en. Retrieved 12 November 2021.
  2. ^ "Appendices | CITES". cites.org. Retrieved 2022-01-14.
  3. ^ a b "Fossa fossana". ITIS. Retrieved 1 January 2015.
  4. ^ Hawkins, A. F .A. (2008). "Fossa fossana". IUCN Red List of Threatened Species. 2008. Retrieved 22 March 2009.{{cite iucn}}: old-form url (help)
  5. ^ Nick Garbutt; Hilary Bradt; Derek Schuurman (2008). Madagascar Wildlife. Bradt Travel Guides. p. 78. ISBN 978-1-84162-245-3.
  6. ^ Wozencraft, W. C. (2005). "Order Carnivora". In Wilson, D. E.; Reeder, D. M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference (3rd ed.). Johns Hopkins University Press. p. 560. ISBN 978-0-8018-8221-0. OCLC 62265494.
  7. ^ Blench, Roger M.; Walsh, Martin (2009). Faunal names in Malagasy: their etymologies and implications for the prehistory of the East African coast (PDF). Eleventh International Conference on Austronesian Linguistics (11 ICAL). Aussois, France. pp. 1–31.
  8. ^ Anjali Goswami; Anthony Friscia (29 July 2010). Carnivoran Evolution: New Views on Phylogeny, Form and Function. Cambridge University Press. pp. 68–70. ISBN 978-1-139-48853-2.
  9. ^ a b "Fossa fossana". animaldiversity.org. Retrieved 1 January 2015.
  10. ^ Steven Roger Fischer (15 February 2013). Islands: From Atlantis to Zanzibar. Reaktion Books. p. 66. ISBN 978-1-78023-053-5.
  11. ^ a b c Nick Garbutt (2007). Mammals of Madagascar: A Complete Guide. Yale University Press. p. 209. ISBN 978-0-300-12550-4.
  12. ^ Richard T. Corlett; Richard B. Primack (3 March 2011). Tropical Rain Forests: An Ecological and Biogeographical Comparison. John Wiley & Sons. p. 136. ISBN 978-1-4443-9228-9.
  13. ^ "Fossa fossana (Malagasy civet)". Animal Diversity Web.
  14. ^ Hilary Bradt; Daniel Austin (1 July 2014). Madagascar. Bradt Travel Guides. p. 66. ISBN 978-1-84162-498-3.
  15. ^ Daniel Austin (10 November 2014). Madagascar Wildlife. Bradt Travel Guides. p. 84. ISBN 978-1-84162-557-7.
  16. ^ Animals: A Visual Encyclopedia (Second Edition): A Visual Encyclopedia. DK Publishing. 19 March 2012. p. 83. ISBN 978-0-7566-9896-6.
  17. ^ "Fossa fossana". Encyclopedia of Life. Retrieved 1 January 2015.
  18. ^ a b Ronald M. Nowak (2005). Walker's Carnivores of the World. JHU Press. pp. 201–202. ISBN 978-0-8018-8032-2.

Sources

  • Macdonald, David (ed). The Encyclopedia of Mammals. (New York, 1984)
  • Anderson, Simon (ed). Simon & Schuster's Guide to Mammals. (Milan, 1982)
Wikispecies has information related to Fossa fossana.
Wikimedia Commons has media related to Fossa_fossana.
  • ARKive - images and movies of the striped civet (Fossa fossana)
  • v
  • t
  • e
Extant Carnivora species
Feliformia ("cat-like" carnivorans)
  • African palm civet (Nandinia binotata)
Prionodon (Asiatic linsangs)
  • Banded linsang (P. linsang)
  • Spotted linsang (P. pardicolor)
Felidae (cats)
Pantherinae
Neofelis
  • Sunda clouded leopard (N. diardi)
  • Clouded leopard (N. nebulosa)
Panthera
  • Snow leopard (P. uncia)
  • Tiger (P. tigris)
  • Jaguar (P. onca)
  • Lion (P. leo)
  • Leopard (P. pardus)
Felinae sensu stricto
Bay cat
lineage
Pardofelis
  • Marbled cat (P. marmorata)
Catopuma
  • Bay cat (C. badia)
  • Asian golden cat (C. temminckii)
Caracal
lineage
  • Serval (Leptailurus serval)
Caracal
  • African golden cat (C. aurata)
  • Caracal (C. caracal)
Leopardus
  • Pampas cat (L. colocola)
  • Geoffroy's cat (L. geoffroyi)
  • Kodkod (L. guigna)
  • Southern tiger cat (L. guttulus)
  • Andean mountain cat (L. jacobita)
  • Ocelot (L. pardalis)
  • Oncilla (L. tigrinus)
  • Margay (L. wiedii)
Lynx
  • Canada lynx (L. canadensis)
  • Eurasian lynx (L. lynx)
  • Iberian lynx (L. pardinus)
  • Bobcat (L. rufus)
Puma
lineage
Acinonyx
  • Cheetah (A. jubatus)
  • Jaguarundi (Herpailurus yagouaroundi)
Puma
  • Cougar (P. concolor)
Leopard cat
lineage
  • Pallas's cat (Otocolobus manul)
Prionailurus
  • Leopard cat (P. bengalensis)
  • Sunda leopard cat (P. javanensis)
  • Flat-headed cat (P. planiceps)
  • Rusty-spotted cat (P. rubiginosus)
  • Fishing cat (P. viverrinus)
Felis
  • Chinese mountain cat (F. bieti)
  • Domestic cat (F. catus)
  • Jungle cat (F. chaus)
  • African wildcat (F. lybica)
  • Sand cat (F. margarita)
  • Black-footed cat (F. nigripes)
  • European wildcat (F. silvestris)
Viverroidea
    • see below↓
Hemigalinae
  • Owston's palm civet (Chrotogale owstoni)
  • Otter civet (Cynogale bennettii)
  • Hose's palm civet (Diplogale hosei)
  • Banded palm civet (Hemigalus derbyanus)
Paradoxurinae
  • Binturong (Arctictis binturong)
  • Small-toothed palm civet (Arctogalidia trivirgata)
  • Sulawesi palm civet (Macrogalidia musschenbroekii)
  • Masked palm civet (Paguma larvata)
Paradoxurus
  • Asian palm civet (P. hermaphroditus)
  • Brown palm civet (P. jerdoni)
  • Golden palm civet (P. zeylonensis)

Viverrinae
sensu lato
Viverrinae
sensu stricto
  • Small Indian civet (Viverricula indica)
  • African civet (Civettictis civetta)
Viverra
  • Malabar large-spotted civet (V. civettina)
  • Large-spotted civet (V. megaspila)
  • Malayan civet (V. tangalunga)
  • Large Indian civet (V. zibetha)
Genettinae
Poiana
(African linsangs)
  • Central African oyan (P. richardsonii)
  • West African oyan (P. leightoni)
Genetta
(genets)
  • Abyssinian genet (G. abyssinica)
  • Angolan genet (G. angolensis)
  • Bourlon's genet (G. bourloni)
  • Crested servaline genet (G. cristata)
  • Common genet (G. genetta)
  • Johnston's genet (G. johnstoni)
  • Letaba genet (G. letabae)
  • Rusty-spotted genet (G. maculata)
  • Pardine genet (G. pardina)
  • Aquatic genet (G. piscivora)
  • King genet (G. poensis)
  • Servaline genet (G. servalina)
  • Hausa genet (G. thierryi)
  • Cape genet (G. tigrina)
  • Giant forest genet (G. victoriae)
  • South African small-spotted genet (G. felina)
Herpestoidea
    • see below↓
Hyaenidae
(hyenas)
Proteles
  • Aardwolf (P. cristata)
Hyaeninae
(bone-crushing hyenas)
  • Striped hyena (Hyaena hyaena)
  • Brown hyena (Parahyaena brunnea)
Crocuta
  • Spotted hyena (C. crocuta)
Herpestidae sensu lato
Eupleridae
(Malagasy
carnivorans)
Euplerinae
  • Fossa (Cryptoprocta ferox)
  • Malagasy civet (Fossa fossana)
Eupleres
  • Eastern falanouc (E. goudotii)
  • Western falanouc (E. major)
Galidiinae
  • Ring-tailed vontsira (Galidia elegans)
Galidictis
  • Broad-striped Malagasy mongoose (G. fasciata)
  • Grandidier's mongoose (G. grandidieri)
  • Narrow-striped mongoose (Mungotictis decemlineata)
Salanoia
  • Brown-tailed mongoose (S. concolor)
  • Durrell's vontsira (S. durrelli)
Herpestidae
sensu stricto

(mongooses)
Mungotinae
Suricata
  • Meerkat (S. suricatta)
  • Liberian mongoose (Liberiictus kuhni)
Mungos
  • Gambian mongoose (M. gambianus)
  • Banded mongoose (M. mungo)
  • Pousargues's mongoose (Dologale dybowskii)
Helogale
  • Ethiopian dwarf mongoose (H. hirtula)
  • Common dwarf mongoose (H. parvula)
Crossarchus
  • Alexander's kusimanse (C. alexandri)
  • Angolan kusimanse (C. ansorgei)
  • Common kusimanse (C. obscurus)
  • Flat-headed kusimanse (C. platycephalus)
Herpestinae
  • Marsh mongoose (Atilax paludinosus)
  • Long-nosed mongoose (Xenogale naso)
Urva
  • Small Indian mongoose (U. auropunctata)
  • Short-tailed mongoose (U. brachyura)
  • Indian grey mongoose (U. edwardsii)
  • Indian brown mongoose (U. fusca)
  • Javan mongoose (U. javanica)
  • Collared mongoose (U. semitorquata)
  • Ruddy mongoose (U. smithii)
  • Crab-eating mongoose (U. urva)
  • Stripe-necked mongoose (U. vitticolla)
  • White-tailed mongoose (Ichneumia albicauda)
  • Yellow mongoose (Cynictis penicillata)
  • Selous's mongoose (Paracynictis selousi)
  • Meller's mongoose (Rhynchogale melleri)
Bdeogale
  • Bushy-tailed mongoose (B. crassicauda)
  • Jackson's mongoose (B. jacksoni)
  • Black-footed mongoose (B. nigripes)
Herpestes
  • Angolan slender mongoose (H. flavescens)
  • Egyptian mongoose (H. ichneumon)
  • Somalian slender mongoose (H. ochracea)
  • Cape gray mongoose (H. pulverulenta)
  • Common slender mongoose (H. sanguinea)
Caniformia ("dog-like" carnivorans)
Canidae (dogs)
Urocyon
  • Gray fox (U. cinereoargenteus)
  • Island fox (U. littoralis)
Vulpini
  • Bat-eared fox (Otocyon megalotis)
Nyctereutes
  • Common raccoon dog (N. procyonoides)
  • Japanese raccoon dog (N. viverrinus)
Vulpes
(foxes)
  • Bengal fox (V. bengalensis)
  • Blanford's fox (V. cana)
  • Cape fox (V. chama)
  • Corsac fox (V. corsac)
  • Tibetan fox (V. ferrilata)
  • Arctic fox (V. lagopus)
  • Kit fox (V. macrotis)
  • Pale fox (V. pallida)
  • Rüppell's fox (V. rueppelli)
  • Swift fox (V. velox)
  • Red fox (V. vulpes)
  • Fennec fox (V. zerda)
Canini
(true dogs)
Cerdocyonina
(zorro)
  • Maned wolf (Chrysocyon brachyurus)
Speothos
  • Bush dog (S. venaticus)
  • Short-eared dog (Atelocynus microtis)
  • Crab-eating fox (Cerdocyon thous)
Lycalopex
  • Culpeo (L. culpaeus)
  • Darwin's fox (L. fulvipes)
  • South American gray fox (L. griseus)
  • Pampas fox (L. gymnocercus)
  • Sechuran fox (L. sechurae)
  • Hoary fox (L. vetulus)
Canina
(wolf-like canids)
Lupulella
  • Side-striped jackal (L. adustus)
  • Black-backed jackal (L. mesomelas)
  • African wild dog (Lycaon pictus)
  • Dhole (Cuon alpinus)
Canis
  • Golden jackal (C. aureus)
  • Domestic dog (C. familiaris)
  • Coyote (C. latrans)
  • African wolf (C. lupaster)
  • Wolf (C. lupus)
  • Eastern wolf (C. lycaon)
  • Red wolf (C. rufus)
  • Ethiopian wolf (C. simensis)
Ursidae
(bears)
Ailuropoda
  • Giant panda (A. melanoleuca)
Tremarctos
  • Spectacled bear (T. ornatus)
Ursinae
  • Sloth bear (Melursus ursinus)
  • Sun bear (Helarctos malayanus)
Ursus
  • American black bear (U. americanus)
  • Asian black bear (U. thibetanus)
  • Brown bear (U. arctos)
  • Polar bear (U. maritimus)
Mustelida
Pinnipedia (seals)
    • see below↓
Musteloidea
    • see below↓
Pinnipedia (seals)
Odobenidae
  • Walrus (Odobenus rosmarus)

Otariidae
(eared seals:
fur seals,
sea lions)
Callorhinus
  • Northern fur seal (C. ursinus)
Otariinae
  • Steller sea lion (Eumetopias jubatus)
Zalophus
  • California sea lion (Z. californianus)
  • Galápagos sea lion (Z. wollebaeki)
  • South American sea lion (Otaria flavescens)
Neophoca
  • Australian sea lion (N. cinerea)
  • New Zealand sea lion (Phocarctos hookeri)
Arctocephalus
  • South American fur seal (A. australis)
  • Australasian fur seal (A. forsteri)
  • Galápagos fur seal (A. galapagoensis)
  • Antarctic fur seal (A. gazella)
  • Juan Fernández fur seal (A. philippii)
  • Brown fur seal (A. pusillus)
  • Guadalupe fur seal (A. townsendi)
  • Subantarctic fur seal (A. tropicalis)
Phocidae
(earless seals
or true seals)
Phocinae
("northern seals")
  • Bearded seal (Erignathus barbatus)
  • Hooded seal (Cystophora cristata)
Phocini
  • Harp seal (Pagophilus groenlandicus)
  • Ribbon seal (Histriophoca fasciata)
  • Grey seal (Halichoerus grypus)
Phoca
  • Spotted seal (P. largha)
  • Harbor seal (P. vitulina)
Pusa
  • Caspian seal (P. caspica)
  • Ringed seal (P. hispida)
  • Baikal seal (P. sibirica)
Monachinae
("southern seals")
Monachini
(monk seals)
  • Mediterranean monk seal (Monachus monachus)
Neomonachus
  • Hawaiian monk seal (N. schauinslandi)
Mirounga
(elephant seals)
  • Northern elephant seal (M. angustirostris)
  • Southern elephant seal (M. leonina)
Lobodontini
(Antarctic seals)
  • Leopard seal (Hydrurga leptonyx)
  • Weddell seal (Leptonychotes weddellii)
  • Crabeater seal (Lobodon carcinophagus)
  • Ross seal (Ommatophoca rossi)
Ailuridae
  • Red panda (Ailurus fulgens)
Mephitidae
(skunks)
Conepatus
(hog-nosed skunks)
  • Molina's hog-nosed skunk (C. chinga)
  • Humboldt's hog-nosed skunk (C. humboldtii)
  • American hog-nosed skunk (C. leuconotus)
  • Striped hog-nosed skunk (C. semistriatus)
Mephitis
  • Hooded skunk (M. macroura)
  • Striped skunk (M. mephitis)
Mydaus
(stink badgers)
  • Sunda stink badger (M. javanensis)
  • Palawan stink badger (M. marchei)
Spilogale
(spotted skunks)
  • Southern spotted skunk (S. angustifrons)
  • Western spotted skunk (S. gracilis)
  • Eastern spotted skunk (S. putorius)
  • Pygmy spotted skunk (S. pygmaea)
Procyonidae
  • Kinkajou (Potos flavus)
Bassariscus
  • Ringtail (B. astutus)
  • Cacomistle (B. sumichrasti)
Procyon
(raccoons)
  • Crab-eating raccoon (P. cancrivorus)
  • Raccoon (P. lotor)
  • Cozumel raccoon (P. pygmaeus)
Bassaricyon
(olingos)
  • Eastern lowland olingo (B. alleni)
  • Northern olingo (B. gabbii)
  • Western lowland olingo (B. medius)
  • Olinguito (B. neblina)
Nasuina
(coatis)
Nasua
  • South American coati (N. nasua)
  • White-nosed coati (N. narica)
Nasuella
  • Eastern mountain coati (N. meridensis)
  • Western mountain coati (N. olivacea)
Mustelidae
    • see below↓
  • American badger (Taxidea taxus)
Mellivora
  • Honey badger (M. capensis)
Melinae
(Eurasian badgers)
Arctonyx
  • Northern hog badger (A. albogularis)
  • Greater hog badger (A. collaris)
  • Sumatran hog badger (A. hoevenii)
Meles
  • Japanese badger (M. anakuma)
  • Caucasian badger (M. canescens)
  • Asian badger (M. leucurus)
  • European badger (M. meles)
Melogale
(ferret-badgers)
  • Vietnam ferret-badger (M. cucphuongensis)
  • Bornean ferret badger (M. everetti)
  • Chinese ferret-badger (M. moschata)
  • Javan ferret-badger (M. orientalis)
  • Burmese ferret-badger (M. personata)
  • Formosan ferret-badger (M. subaurantiaca)
Guloninae
  • Tayra (Eira barbara)
Pekania
  • Fisher (P. pennanti)
Gulo
  • Wolverine (G. gulo)
Martes
(martens)
  • American marten (M. americana)
  • Pacific marten (M. caurina)
  • Yellow-throated marten (M. flavigula)
  • Beech marten (M. foina)
  • Nilgiri marten (M. gwatkinsii)
  • European pine marten (M. martes)
  • Japanese marten (M. melampus)
  • Sable (M. zibellina)
Ictonychinae
Lyncodontini
  • Patagonian weasel (Lyncodon patagonicus)
Galictis
(grisons)
  • Lesser grison (G. cuja)
  • Greater grison (G. vittata)
Ictonychini
(African polecats)
Vormela
  • Marbled polecat (V. peregusna)
  • African striped weasel (Poecilogale albinucha)
Ictonyx
  • Saharan striped polecat (I. libycus)
  • Striped polecat (I. striatus)
Lutrinae
(otters)
  • Giant otter (Pteronura brasiliensis)
Lontra
  • North American river otter (L. canadensis)
  • Marine otter (L. felina)
  • Neotropical otter (L. longicaudis)
  • Southern river otter (L. provocax)
Enhydra
  • Sea otter (E. lutris)
  • Spotted-necked otter (Hydrictis maculicollis)
Lutra
  • Eurasian otter (L. lutra)
  • Hairy-nosed otter (L. sumatrana)
Lutrogale
  • Smooth-coated otter (L. perspicillata)
Aonyx
  • African clawless otter (A. capensis)
  • Asian small-clawed otter (A. cinereus)
  • Congo clawless otter (A. congicus)
Mustelinae
Neogale
  • Amazon weasel (N. africana)
  • Colombian weasel (N. felipei)
  • Long-tailed weasel (N. frenata)
  • American mink (N. vison)
Mustela
(weasels)
  • Sichuan weasel (M. aistoodonnivalis)
  • Mountain weasel (M. altaica)
  • Stoat/Beringian ermine (M. erminea)
  • Steppe polecat (M. eversmannii)
  • Ferret (M. furo)
  • Haida ermine (M. haidarum)
  • Japanese weasel (M. itatsi)
  • Yellow-bellied weasel (M. kathiah)
  • European mink (M. lutreola)
  • Indonesian mountain weasel (M. lutreolina)
  • Black-footed ferret (M. nigripes)
  • Least weasel (M. nivalis)
  • Malayan weasel (M. nudipes)
  • European polecat (M. putorius)
  • American ermine (M. richardsonii)
  • Siberian weasel (M. sibirica)
  • Back-striped weasel (M. strigidorsa)
  • v
  • t
  • e
Genera of civets, mongooses, hyenas, cats, and their extinct allies
Palaeogalidae
Nimravidae
    • See Nimravidae
Aeluroidea
Nandiniidae
Viverroidea
Viverridae
Genettinae
Hemigalinae
Paradoxurinae
Viverrinae
Herpestoidea
Herpestidae
    • See below↓
Hyaenidae
    • See below↓
Feloidea
    • See below↓
Eupleridae
Euplerinae
Galidiinae
Herpestidae
  • Kichechia
  • Legetetia
  • Leptoplesictis
  • Ugandictis
Herpestinae
Mungotinae
Percrocutidae?
Lophocyonidae
Hyaenidae
  • Allohyaena
  • Belbus
  • Metahyaena
  • Pliocrocuta
  • Tongxinictis
  • Werdelinus
Ictitheriinae
Protelinae
Hyaeninae
Barbourofelidae
  • (See Barbourofelidae)
Prionodontidae
Felidae
Proailurinae
Felinae
Machairodontinae
    • See Machairodontinae
Pantherinae
Taxon identifiers
Fossa fossana