Ensete ventricosum
Ensete ventricosum | |
---|---|
![]() | |
Ensete ventricosum, by Walter Hood Fitch (1861) | |
Conservation status | |
![]() Least Concern (IUCN 3.1)[1] | |
Scientific classification ![]() | |
Kingdom: | Plantae |
Clade: | Tracheophytes |
Clade: | Angiosperms |
Clade: | Monocots |
Clade: | Commelinids |
Order: | Zingiberales |
Family: | Musaceae |
Genus: | Ensete |
Species: | E. ventricosum |
Binomial name | |
Ensete ventricosum (Welw.) Cheesman | |
Synonyms[2] | |
|
Ensete ventricosum, commonly known as enset or ensete, Ethiopian banana, Abyssinian banana,[3] pseudo-banana, false banana and wild banana,[4] is a species of flowering plant in the banana family Musaceae. The domesticated form of the plant is cultivated only in Ethiopia, where it provides the staple food for approximately 20 million people.[5][6] The name Ensete ventricosum was first published in the Kew Bulletin[7] 1947, p. 101. Its synonyms include Musa arnoldiana De Wild., Musa ventricosa Welw. and Musa ensete J. F. Gmelin.[8] In its wild form, it is native to the eastern edge of the Great African Plateau, extending northwards from South Africa through Mozambique, Zimbabwe, Malawi, Kenya, Uganda and Tanzania to Ethiopia, and west to the Congo, being found in high-rainfall forests on mountains, and along forested ravines and streams.[5]
Description
Like bananas, Ensete ventricosum is a large non-woody plant—a gigantic monocarpic evergreen perennial (not a tree)[9]—up to 6 m (20 ft) tall. The tallest to be reported was 42 feet (13 meters).[10] It has a stout pseudostem of tightly overlapping leaf bases, and large banana-like leaf blades of up to 5 m (16 ft) tall by 1 m (3 ft 3 in) wide, with a salmon-pink midrib. A mature plant weighs about 550 pounds (250 kilograms).[11] The flowers, which only occur once from the centre of the plant at the end of that plant's life,(monocarpic) are in massive pendant thyrsesup to ten feet ( three meters) in length bearing thirty or more "hands" of young bananas which are covered by large pink bracts. The roots are an important foodstuff, but the fruits are inedible (insipid, flavorless)[12] and have hard, black, rounded seeds.
The Latin specific epithet ventricosum means “with a swelling on the side, like a belly”.[13]
Use as a foodcrop
![](http://upload.wikimedia.org/wikipedia/commons/thumb/a/af/WomanEnset.jpg/220px-WomanEnset.jpg)
Enset is a very important local food source especially in Ethiopia. The Food and Agriculture Organization of the United Nations reports that "enset provides more amount of foodstuff per unit area than most cereals. It is estimated that 40 to 60 enset plants occupying 250–375 square metres (2,700–4,000 sq ft) can provide enough food for a family of 5 to 6 people."[14][failed verification]
Enset (E. ventricosum) is Ethiopia's most important root crop, a traditional staple in the densely populated south and southwestern parts of Ethiopia.[15] Its importance to the diet and economy of the Gurage and Sidama peoples was first recorded by Jerónimo Lobo in the seventeenth century.[16] Each plant takes four to five years to mature, at which time a single root will yield about 40 kg (88 lb) of food. Because of the long period of time from planting to harvest, plantings need to be staggered over time, to ensure that there is enset available for harvest in every season. Enset will tolerate drought better than most cereal crops.
Wild enset plants are produced from seeds, while most domesticated plants are propagated from suckers. Up to 400 suckers can be produced from just one mother plant. In 1994 3,000 km2 (1,200 sq mi) of enset were grown in Ethiopia, with a harvest estimated to be almost 10 tonnes per hectare (4.0 long ton/acre; 4.5 short ton/acre). Enset is often intercropped with sorghum, although the practice amongst the Gedeo people is to intercrop it with coffee.[17]
The young and tender tissues in the centre or heart of the plant (the growing point) are cooked and eaten, being tasty and nutritious and very like the core of palms and cycads. In Ethiopia, more than 150,000 hectares (370,000 acres; 580 sq mi) are cultivated for the starchy staple food prepared from the pulverised trunk and inflorescence stalk. Fermenting these pulverised parts results in a food called kocho. Bulla is made from the liquid squeezed out of the mixture and sometimes eaten as a porridge, while the remaining solids are suitable for consumption after a settling period of some days. Mixed kocho and bulla can be kneaded into dough, then flattened and baked over a fire. Kocho is in places regarded as a delicacy, suitable for serving at feasts and ceremonies such as weddings, when wheat flour is added. The fresh corm is cooked like potatoes before eating. Dry kocho and bulla are energy-rich and produce from 14 to 20 kJ/g (3.3 to 4.8 kcal/g).
It is a major crop, although often supplemented with cereal crops.[18] However its value as a famine food has fallen for a number of reasons, as detailed in the April 2003 issue of the UN-OCHA Ethiopia unit's Focus on Ethiopia:
Apart from an enset plant disease epidemic in 1984–85 which wiped out large parts of the plantations and created the green famine, in the past 10 years major factors were recurrent drought and food shortage together with acute land shortage that forced farmers more and more into consumption of immature plants. Hence farmers were overexploiting their Enset reserves thereby causing gradual losses and disappearance of the false banana as an important household food security reserve. Even though not all the plant losses can be attributed to drought and land shortage and hence early consumption of immature crops, estimations go as far as more than 60% of the false banana crop stands have been lost in some areas in SNNPR during the last 10 years. This basically means that a great many people who used to close the food gap with false banana consumption are not able to do so any more, and lacking a viable alternative, have become food insecure and highly vulnerable to climatic and economic disruptions of their agricultural system.[19]
Other uses
The plant is quick-growing and often cultivated as an ornamental plant. In frost-prone areas it requires winter protection under glass.[9] It has gained the Royal Horticultural Society's Award of Garden Merit,[20][21] as has the cultivar 'Maurelii' (Ethiopian black banana)[22]
A good quality fibre, suitable for ropes, twine, baskets, and general weaving, is obtained from the leaves. Dried leaf-sheaths are used as packing material, serving the same function as Western foam plastic and polystyrene. The entire plant but the roots is used to feed livestock.[23][24] Fresh leaves are a common fodder for cattle during the dry season,[24] and many farmers feed their animals with residues of enset harvest or processing.[24]
History
In 1769, the celebrated Scottish traveller James Bruce first sent a description and quite accurate drawings of a plant common in the marshes around Gondar in Ethiopia, confidently pronounced it to be "no species of Musa" and wrote that its local name was "ensete". In 1853 the British Consul at Mussowah sent some seeds to Kew Gardens, mentioning that their native name was ansett. Kew, quite understandably, did not make the connection, especially as they had never before seen such seeds. However, when the seeds had germinated and the plants had rapidly gained size, their relationship to the true banana became obvious.
Bruce also discussed the plant's place in the mythology of Egypt and pointed out that some Egyptian statue carvings depict the goddess Isis sitting among the leaves of what was thought to be a banana plant, a plant native to Southeast Asia and not known in Ancient Egypt.[25][26]
Pests and diseases
A major issue with the cultivation of enset is its vulnerability to several pests and diseases.
Pests
The most common pest that threatens enset is caused by the Cataenococcus enset which is a root mealybug. The Cataenococcus enset feeds on the roots and corm of the enset plant which leads to slower growth and easier uprooting. Even though enset can be infested at all age stages, the highest risk is between the second or fourth growth year.[27] The dispersion of the mealybug occurs through different vectors: First, the larvae can crawl short distances[27] as adults mealybugs tend to move only after being disturbed.[28] Second, mealybugs-ant symbiotic relationships can be linked to enset infestation and protect and even transport the mealybug over short distances. In return, they feed on the mealybug honeydew. Third, flooding events can transport the mealybug over longer distances and reach enset plants. However, the main transport vectors are unclean working tools and the usage of already infected suckers.[27] This means that the best way to get rid of the bug and to limit its propagation is to uproot the plant and burn it.[29][30] In addition, the fields can be kept free of plant growth for a month since the mealybug can only survive up to three weeks without plant material.[27][28]
Other pests include nematodes, spider mites, aphids, mole rats, porcupines and wild pigs. The latter erode the corm and pseudostem.[27] As for the nematodes, there are two predominant species: there are the root lesion nematodes (Pratylenchus goodeyi) and the root-knot nematodes (Meloidogyne sp.) and their appearance stand in connection with bacterial wilt.[27] Pratylenchus goodeyi create lesions on the corm and roots, which can lead to cavities up to 2 cm (0.79 in) and characteristic purple colouring around the cavities. The nematode infestation leads to the easy uprooting of the affected plants. Crop rotation can counteract high nematode infestations.[31]
Diseases
![](http://upload.wikimedia.org/wikipedia/commons/thumb/c/c4/Platano_Colombia1_%284875321613%29.jpg/220px-Platano_Colombia1_%284875321613%29.jpg)
The enset plant can be subject to multiple diseases that threaten its use in agriculture.[32][30] The most well known of them is the infection by the bacteria Xanthomonas campestris pathovar Musacerum which creates bacterial wilt, also known as borijje and wol’a by the Kore people.[29] The first observation of this disease was reported by Yirgou and Bradbury in 1968.[33] The manifestation of the bacterial wilt is taking place in the apical leaves that will wilt then dry and finally lead to the drying of the whole plant.[29][34] The only way to avoid the spreading of the disease is in uprooting, burning and burying plants as well as in applying strict control of the knives and tools used to harvest and treat the plants.
Other diseases have been observed such as Okka and Woqa which occur respectively in case of severe drought and in situations of too much water in the soil which causes the proliferation of bacteria. These problems can be solved by either watering the field when drought is present or by draining the soil to avoid too much water.[29]
Another disease can strike enset even though it has been more observed on Banana plants (Musacea). This disease is caused by Mycospharella spp. and is commonly called black Sigatoka leaf streaks. The symptoms are basically dark/brown lesions surrounded by yellow on the leaves.[35][36][37][38] This disease happens to be favoured by high rainfall and lower temperature.[39][40][41]
Socio-cultural importance of enset in Ethiopia
Enset cultivation in Ethiopia is reported to be 10,000 years old, though there is little empirical evidence to support this.[42][43][29] It has major economic, social, cultural and environmental functions related to trade, medicine, cultural identity, rituals or settlement patterns.[42][29][44]
The Enset-planting complex is one of the four farming systems of Ethiopia together with pastoralism, shifting cultivation and the seed-farming complex. It is widely used by around 20 million people, which represent 20-25% of the population. They mainly live in the densely populated highlands of south and southwest Ethiopia.[43][29]
The plant is very important for food security because it is quite resistant to droughts (the growth only stops for a short time) and it can be harvested at any development stage.[43] Nevertheless, in recent years, the population growth has put pressure on enset cultivation systems. This is mainly because of a decrease of fertilization through manure and an increase in demand, especially during droughts. At such times, enset becomes the only resource available.[43]
The role of gender in enset cultivation
Gender roles in enset cultivation are of high importance,[42] as a strong division of work exists: generally men are responsible for the propagation, cultivation, and transplanting of enset, while women are in charge of manuring, hand-weeding, thinning and landrace selection.[29][43] Additionally, women process enset plants, which is a tedious work (transformation of the plant into useful material, principally food and fibres) for which they generally come together. Men are banned from the field during this process.[42][43][29] As women are responsible to provide sufficient food to their family, they are the ones who choose when and which plant to harvest and which quantity to sell.[32]
Several studies state the importance of women's knowledge on the different crop varieties. Women are more likely to precisely recognize the different varieties of the plant than men.[42][32][43] Nevertheless, women's work is often neglected or considered of lesser importance than men's by researchers and farmers[42] and women are less likely to get access to extension services and quality services than men.[45]
Another important aspect in which gender plays a role is in the classification of enset varieties. Indeed, they differentiate "male" varieties from "female" varieties, according to the preferences of men and women who harvest them.[32] Whereas men prefer late maturing genotypes resistant to diseases, women prefer varieties that are good for cooking and can be harvested for consumption at an earlier stage.[32] In general, households tend to have slightly more “female” genotypes than "male" ones.[42][32]
Enset biodiversity and socio-cultural and -economic groups
In Ethiopia, over 300 enset varieties have been recorded[46] which is important for agro- and biodiversity. The farmers’ main interest for maintaining biodiversity is the different beneficial characteristics of each variety.[47] This means that Ethiopian farmers spread important characteristics over many enset varieties instead of combining a number of desired characteristics in one single genotype[47] This is a significant difference between Ethiopian subsistence farmers’ and plant breeders’ approaches.
More than 11 ethnic groups with different cultures, traditions and agricultural systems inhabit the enset-growing regions. This contributes to the high number of varieties.[43] Over centuries, the different ethnic groups have applied their specific indigenous knowledge of farming systems in order to sustain production in various ways. A dying out of enset varieties would hence also make disappear a part of cultural practices and linguistic terms in Ethiopia (Negash et al., 2004).[47]
Enset biodiversity is preserved not only due to the presence of different ethnic groups but also due to different households’ wealth status. Richer farmers can generally afford to maintain a higher level of farm biodiversity because they have more resources such as land, labour and livestock. Therefore, they can cultivate more varieties with differing specific characteristics.[43] However, also poorer households try to maintain as many clones as possible by selecting the disease-resistant first.[47]
Known variants and hybrids
- Ensete ventricosum 'Atropurpureum'
- Ensete ventricosum 'Green Stripe'
- Red false banana (Ensete ventricosum 'Maurelii', syn. Musa maurelii)
- Ensete ventricosum 'Hiniba'[48]
- Ensete ventricosum 'Montbeliardii'
- Ensete ventricosum 'Tandarra Red' (syn. Musa 'Tandarra Red')
- Ensete ventricosum 'Red Stripe' (syn. Musa 'Red Stripe')
- Ensete ventricosum 'Rubra' (syn. Musa ensete 'Rubra')
Sir John Kirk felt that in habit Ensete livingstonianum was indistinguishable from E. ventricosum and noted that both are found in the mountains of equatorial Africa.
Gallery
- Specimen in Jardin botanique exotique de Menton, Alpes-Maritimes, France
- Flower detail
- Plant on Mount Tsetserra, Mozambique
- Stem detail
References
- ^ Williams, E. (2017). "Ensete ventricosum". IUCN Red List of Threatened Species. 2017: e.T22486245A22486942. doi:10.2305/IUCN.UK.2017-3.RLTS.T22486245A22486942.en. Retrieved 21 January 2022.
- ^ "The Plant List: A Working List of all Plant Species".
- ^ "Ensete ventricosum". Germplasm Resources Information Network. Agricultural Research Service, United States Department of Agriculture. Retrieved 20 December 2017.
- ^ "Ensete ventricosum". PlantZAfrica. South African National Biodiversity Institute (SANBI). Retrieved 4 December 2022.
- ^ a b Wilkin, Paul; Demissew, Sebsebe; Willis, Kathy; Woldeyes, Feleke; Davis, Aaron P.; Molla, Ermias L.; Janssens, Steven; Kallow, Simon; Berhanu, Admas (2019). "Enset in Ethiopia: a poorly characterized but resilient starch staple". Annals of Botany. 123 (5): 747–766. doi:10.1093/aob/mcy214. PMC 6526316. PMID 30715125.
- ^ "A 'banana falsa' que pode ser solução para alimentar milhões". BBC News Brasil.
- ^ Cheesman, E. E. (1947). "Classification of the bananas". Kew Bulletin. 2 (2): 97–106. doi:10.2307/4109206. JSTOR 4109206. Retrieved 22 January 2022.
- ^ Wikipedia DE[circular reference]
- ^ a b RHS A-Z encyclopedia of garden plants. United Kingdom: Dorling Kindersley. 2008. p. 1136. ISBN 978-1-4053-3296-5.
- ^ Brenan, J.P.N.; Greenway, P.J. (1949). Checklists of the Forest Trees and Shrubs of the British Empire - #5 - Tanganyika Territory - Part 2. Oxford, England: Imperial Forestry Institute. p. 364.
- ^ gardener's chronicle Volume 27 (third series) February 3, 1900 Page 69. Quoting Kew Bull
- ^ "Ensete ventricosum". ecocrop.fao.org. Archived from the original on 30 June 2018.
- ^ Harrison, Lorraine (2012). RHS Latin for Gardeners. United Kingdom: Mitchell Beazley. ISBN 978-1845337315.
- ^ Country Information Brief, FAO June 1995.
- ^ Richard Pankhurst, Economic History of Ethiopia (Addis Ababa: Haile Selassie I University, 1968), p. 194. Pankhurst uses the taxononym Musa ensete.
- ^ Jerónimo Lobo, The Itinerário of Jerónimo Lobo, translated by Donald M. Lockhart (London: Hakluyt Society, 1984), pp. 245f
- ^ Kippie Kanshie, T. "Five thousand years of sustainability? A case study on Gedeo land use" Archived 28 September 2011 at the Wayback Machine (PhD dissertation: May 2002), p. 38
- ^ Kippie Kanshie, T. "Five thousand years", p. 19
- ^ "Enset as staple food not valuable anymore to bridge food gap", Focus on Ethiopia, April 2003, UN-OCHA-Ethiopia (accessed 3 March 2009)
- ^ "Ensete ventricosum". Royal Horticultural Society. Retrieved 7 June 2020.
- ^ "AGM Plants - Ornamental" (PDF). Royal Horticultural Society. July 2017. p. 35. Retrieved 6 February 2018.
- ^ "Ensete ventricosum 'Maurelii'". RHS. Retrieved 7 June 2020.
- ^ "Plant Resources of Tropical Africa".
- ^ a b c Heuzé V., Thiollet H., Tran G., Hassoun P., Lebas F., 2016. Enset (Ensete ventricosum) corms and pseudostems. Feedipedia, a programme by INRA, CIRAD, AFZ and FAO. https://www.feedipedia.org/node/21251
- ^ Curtis's Botanical Magazine vol. 87 (1861)
- ^ Macfarquhar, Colin; Gleig, George (4 June 1797). Encyclopædia Britannica: Or, A Dictionary of Arts, Sciences, and Miscellaneous Literature. A. Bell and C. Macfarquhar. p. 469 – via Internet Archive.
Maitsha Ethiopia.
- ^ a b c d e f P., Parvatha Reddy (5 June 2015). Plant protection in tropical root and tuber crops. New Delhi. ISBN 9788132223894. OCLC 910878064.
{{cite book}}
: CS1 maint: location missing publisher (link) - ^ a b Addis, Temesgen; Azerefegne, Ferdu; Blomme, Guy; Kanaujia, K (2008). "Biology of the Enset Root Mealybug Cataenococcus ensete and its Geographical Distribution in Southern Ethiopia". Journal of Applied Biosciences. 8 (1): 251–260 – via Researchgate.
- ^ a b c d e f g h i Assoma, Awoke Amzaye; Hewlett, Barry S. Adaptation and Change in Enset Ecology and Farming Among the Kore of Southwestern Ethiopia. OCLC 1056710194.
- ^ a b Addis, T; Azerefegne, F; Blomme, G (11 May 2010). "Density and distribution on enset root mealybugs on enset". African Crop Science Journal. 16 (1). doi:10.4314/acsj.v16i1.54344. hdl:1807/47442.
- ^ Peregrine, W. T. H.; Bridge, John (January 1992). "The lesion nematode Pratylenchus goodeyian important pest of Ensete in Ethiopia". Tropical Pest Management. 38 (3): 325–326. doi:10.1080/09670879209371719.
- ^ a b c d e f Negash, Almaz; Niehof, Anke (2004). "The significance of enset culture and biodiversity for rural household food and livelihood security in southwestern Ethiopia". Agriculture and Human Values. 21 (1): 61–71. doi:10.1023/b:ahum.0000014023.30611.ad. S2CID 153467789.
- ^ Yirgou, D.; Bradbury, J. F. (19 March 1974). "A Note on Wilt of Banana Caused by the Enset Wilt Organism Xanthomonas musacearum". East African Agricultural and Forestry Journal. 40 (1): 111–114. doi:10.1080/00128325.1974.11662720.
- ^ Nakato, Valentine; Mahuku, George; Coutinho, Teresa (20 September 2017). "Xanthomonas campestris pv. musacearum: a major constraint to banana, plantain and enset production in central and east Africa over the past decade". Molecular Plant Pathology. 19 (3): 525–536. doi:10.1111/mpp.12578. PMC 6638165. PMID 28677256.
- ^ Gurmu, Tadesse; Adugna, Girma; Berecha, Gezahegn (28 December 2016). "Black Sigatoka leaf streaks of banana (Musa spp.) caused by Mycosphaerella fijiensis in Ethiopia". Journal of Plant Diseases and Protection. 124 (3): 245–253. doi:10.1007/s41348-016-0070-8. S2CID 91077470.
- ^ Jones, David Robert (1999). "Black leaf streak: Symptoms". Diseases of banana, abacá, and enset. Wallingford, Oxon, UK: CABI Publishing. pp. 44–48. ISBN 978-0851993553. OCLC 41347037.
- ^ Marín, Douglas H.; Romero, Ronald A.; Guzmán, Mauricio; Sutton, Turner B. (2003). "Black Sigatoka: An Increasing Threat to Banana Cultivation". Plant Disease. 87 (3). American Phytopathological Society (APS): 208–222. doi:10.1094/pdis.2003.87.3.208. ISSN 0191-2917. PMID 30812750.
- ^ Ministry of Agriculture and Rural Development (MoRA). (2009). Crop variety register issueNo. 12. pp. 157–159.
- ^ Swennen, R.; Vuylsteke, D. (1993). "Breeding Black Sigatoka resistant plantains with a wild banana". Tropical Agriculture. 70 (1): 74–77.
- ^ Tushemereirwe, W. K.; Waller, J. M. (1993). "Black leaf streak (Mycosphaerella fijiensis) in Uganda". Plant Pathology. 42 (3): 471–472. doi:10.1111/j.1365-3059.1993.tb01525.x.
- ^ Zandjanakou-Tachin, M.; Ojiambo, P. S.; Vroh-Bi, I.; Tenkouano, A.; Gumedzoe, Y. M.; Bandyopadhyay, R. (2 July 2012). "Pathogenic variation of Mycosphaerella species infecting banana and plantain in Nigeria". Plant Pathology. 62 (2). American Phytopathological Society (APS): 298–308. doi:10.1111/j.1365-3059.2012.02650.x.
- ^ a b c d e f g Brandt, S. A., Spring, A., Hiebsch, C., McCabe, J. T., Tabogie, E., Diro, M., … Tesfaye, S. (1997). The “Tree Against Hunger,” 66.
- ^ a b c d e f g h i Tsegaye, Admasu (2002). On indigenous production, genetic diversity and crop ecology of enset (Ensete ventricosum (Welw.) Cheesman). s.n. ISBN 978-9058086280. OCLC 906993853.
- ^ Borrell, James S.; Goodwin, Mark; Blomme, Guy; Jacobsen, Kim; Wendawek, Abebe M.; Gashu, Dawd; Lulekal, Ermias; Asfaw, Zemede; Demissew, Sebsebe; Wilkin, Paul (2020). "Enset-based agricultural systems in Ethiopia: A systematic review of production trends, agronomy, processing and the wider food security applications of a neglected banana relative". Plants, People, Planet. 2 (3): 212–228. doi:10.1002/ppp3.10084.
- ^ Ragasa, Catherine; Berhane, Guush; Tadesse, Fanaye; Taffesse, Alemayehu Seyoum (October 2013). "Gender Differences in Access to Extension Services and Agricultural Productivity" (PDF). The Journal of Agricultural Education and Extension. 19 (5): 437–468. doi:10.1080/1389224x.2013.817343. S2CID 18286376.
- ^ Yemataw, Zerihun; Tesfaye, Kassahun; Zeberga, Awole; Blomme, Guy (1 September 2016). "Exploiting indigenous knowledge of subsistence farmers' for the management and conservation of Enset (Ensete ventricosum (Welw.) Cheesman) (Musaceae family) diversity on-farm". Journal of Ethnobiology and Ethnomedicine. 12 (1): 34. doi:10.1186/s13002-016-0109-8. PMC 5009499. PMID 27586388.
- ^ a b c d Negash, Almaz; Niehof, Anke (2004). "The significance of enset culture and biodiversity for rural household food and livelihood security in southwestern Ethiopia". Agriculture and Human Values. 21 (1): 61–71. doi:10.1023/b:ahum.0000014023.30611.ad. S2CID 153467789.
- ^ "Ensete ventricosum 'Hiniba' | /RHS Gardening".
External links
![](http://upload.wikimedia.org/wikipedia/en/thumb/4/4a/Commons-logo.svg/30px-Commons-logo.svg.png)
Data related to Ensete ventricosum at Wikispecies
- Dressler, S.; Schmidt, M. & Zizka, G. (2014). "Ensete ventricosum". African plants – a Photo Guide. Frankfurt/Main: Forschungsinstitut Senckenberg.